Profile

Dr Richard Fox

ESI Affiliate

rf382@exeter.ac.uk

01626 368385

Overview

I am a professional ecologist working in biodiversity conservation and a long-time butterfly and moth enthusiast (on twitter as @RichardFoxBC). I have worked for the UK charity Butterfly Conservation (www.butterfly-conservation.org) since 1997, leading their work on butterfly and moth recording schemes and citizen science. I have been involved in a large number of research collaborations utilising the data gathered through these schemes and recently completed a PhD by Publication at the Environment and Sustainability Institute, Exeter University (Penryn).

Broad research specialisms:

Insect ecology
Biodiversity conservation
Impacts of global environmental change on biodiversity
Citizen science

Qualifications

2020 PhD by Publication Biosciences, University of Exeter

1992 MSc Nature Conservation, University College London

1990 BA Zoology, University of Oxford

Research

Research projects

Measuring UK trends in Lepidopteran biodiversity and assessing extinction risk, and relating these to global efforts to understand biodiversity decline. Ongoing long-term work utilising citizen science gathered data on the occurrence or abundance of butterflies and moths. Current focus is on a new State of Britain's Moths report.
Assessing the impacts of climate change on UK butterflies and moths. Partner in two recent NERC research projects led by the University of York: The velocity of evolutionary responses of species to ecological change; testing adaptive limits in time and space and Refugial populations at trailing-edge range margins: attributes, survival and conservation.
Impact of artificial light at night (ALAN) on moth populations and related ecosystem functioning. Current project is a PhD studentship in collaboration with Newcastle University and the UK Centre for Ecology & Hydrology investigating the impact of ALAN on the abundance and behaviour of moth larvae and cascading effects among their parasitoids communities.
Understanding the benefits and limitations of citizen science. Recent innovative projects have assessed the efficacy of citizen science, patterns of engagement and implications for data quality and the onward use of data contributed by citizens, so as to maximise the benefits of such schemes for participants, researchers and wider society.

Publications

Journal articles

Montràs-Janer T, Suggitt AJ, Fox R, Jönsson M, Martay B, Roy DB, Walker KJ, Auffret AG (2024). Anthropogenic climate and land-use change drive short- and long-term biodiversity shifts across taxa. Nat Ecol Evol, 8(4), 739-751. Abstract. Author URL.

Montràs-Janer T, Suggitt AJ, Fox R, Jönsson M, Martay B, Roy DB, Walker KJ, Auffret AG (2024). Author Correction: Anthropogenic climate and land-use change drive short- and long-term biodiversity shifts across taxa. Nat Ecol Evol, 8(4). Author URL.

Hickinbotham EJ, Pattison Z, Fox R, Rushton SP (2024). Drivers of moth phenology in England and Wales. JOURNAL OF INSECT CONSERVATION Author URL.

Suggitt AJ, Wheatley CJ, Aucott P, Beale CM, Fox R, Hill JK, Isaac NJB, Martay B, Southall H, Thomas CD, et al (2023). Linking climate warming and land conversion to species' range changes across Great Britain. Nat Commun, 14(1). Abstract. Author URL.

Hordley LA, Fox R, Suggitt AJ, Bourn NAD (2023). Precipitation buffers temperature-driven local extinctions of moths at warm range margins. Ecol Lett, 26(5), 805-815. Abstract. Author URL.

Pocock MJO, Logie M, Isaac NJB, Fox R, August T (2023). The recording behaviour of field-based citizen scientists and its impact on biodiversity trend analysis. ECOLOGICAL INDICATORS, 151 Author URL.

Fox R, Dennis EB, Brown AF, Curson J (2022). A revised Red List of British butterflies. INSECT CONSERVATION AND DIVERSITY, 15(5), 485-495. Author URL.

Tordoff GM, Dennis EB, Fox R, Cook PM, Davis TM, Blumgart D, Bourn NAD (2022). Inconsistent results from trait-based analyses of moth trends point to complex drivers of change. BIODIVERSITY AND CONSERVATION, 31(12), 2999-3018. Author URL.

Cook PM, Tordoff GM, Davis TM, Parsons MS, Dennis EB, Fox R, Botham MS, Bourn NAD (2022). Traits data for the butterflies and macro-moths of Great Britain and Ireland. Ecology, 103(5). Abstract. Author URL.

Wagner DL, Fox R, Salcido DM, Dyer LA (2021). A window to the world of global insect declines: Moth biodiversity trends are complex and heterogeneous. Proc Natl Acad Sci U S A, 118(2). Abstract. Author URL.

Sánchez-Fernández D, Fox R, Dennis RLH, Lobo JM (2021). How complete are insect inventories? an assessment of the british butterfly database highlighting the influence of dynamic distribution shifts on sampling completeness. Biodivers Conserv., 30(3), 889-902. Abstract.

Wilson RJ, Fox R (2021). Insect responses to global change offer signposts for biodiversity and conservation. ECOLOGICAL ENTOMOLOGY, 46(4), 699-717. Author URL.

Boyes DH, Evans DM, Fox R, Parsons MS, Pocock MJO (2021). Is light pollution driving moth population declines? a review of causal mechanisms across the life cycle. INSECT CONSERVATION AND DIVERSITY, 14(2), 167-187. Author URL.

Lobo JM, Gutierrez-Canovas C, Fox R, Dennis RLH, Sanchez-Fernandez D (2021). Species traits influence the process of biodiversity inventorying: a case study using the British butterfly database. INSECT CONSERVATION AND DIVERSITY, 14(6), 748-755. Author URL.

Boyes DH, Evans DM, Fox R, Parsons MS, Pocock MJO (2021). Street lighting has detrimental impacts on local insect populations. Sci Adv, 7(35). Abstract. Author URL.

August T, Fox R, Roy DB, Pocock MJO (2020). Data-derived metrics describing the behaviour of field-based citizen scientists provide insights for project design and modelling bias. Sci Rep, 10(1). Abstract. Author URL.

Montgomery GA, Dunn RR, Fox R, Jongejans E, Leather SR, Saunders ME, Shortall CR, Tingley MW, Wagner DL (2020). Is the insect apocalypse upon us? How to find out. Biological Conservation, 241, 108327-108327.

Norberg A, Abrego N, Blanchet FG, Adler FR, Anderson BJ, Anttila J, Araújo MB, Dallas T, Dunson D, Elith J, et al (2019). A comprehensive evaluation of predictive performance of 33 species distribution models at species and community levels. Ecological Monographs, 89(3). Abstract.

Outhwaite CL, Powney GD, August TA, Chandler RE, Rorke S, Pescott OL, Harvey M, Roy HE, Fox R, Roy DB, et al (2019). Annual estimates of occupancy for bryophytes, lichens and invertebrates in the UK, 1970–2015. Scientific Data, 6(1). Abstract.

Boyes DH, Fox R, Shortall CR, Whittaker RJ (2019). Bucking the trend: the diversity of Anthropocene ‘winners’ among British moths. Frontiers of Biogeography, 11(3).

Macgregor CJ, Thomas CD, Roy DB, Beaumont MA, Bell JR, Brereton T, Bridle JR, Dytham C, Fox R, Gotthard K, et al (2019). Climate-induced phenology shifts linked to range expansions in species with multiple reproductive cycles per year. NATURE COMMUNICATIONS, 10 Author URL.

Macgregor CJ, Kitson JJN, Fox R, Hahn C, Lunt DH, Pocock MJO, Evans DM (2019). Construction, validation, and application of nocturnal pollen transport networks in an agro-ecosystem: a comparison using light microscopy and DNA metabarcoding. ECOLOGICAL ENTOMOLOGY, 44(1), 17-29. Author URL.

Macgregor CJ, Pocock MJO, Fox R, Evans DM (2019). Effects of street lighting technologies on the success and quality of pollination in a nocturnally pollinated plant. Ecosphere, 10(1). Abstract.

Dennis EB, Morgan BJT, Fox R, Roy DB, Brereton TM (2019). Functional data analysis of multi-species abundance and occupancy data sets. Ecological Indicators, 104, 156-165. Abstract.

Platts PJ, Mason SC, Palmer G, Hill JK, Oliver TH, Powney GD, Fox R, Thomas CD (2019). Habitat availability explains variation in climate-driven range shifts across multiple taxonomic groups. Scientific Reports, 9(1). Abstract.

Fox R, Harrower CA, Bell JR, Shortall CR, Middlebrook I, Wilson RJ (2019). Insect population trends and the IUCN Red List process. Journal of Insect Conservation, 23(2), 269-278. Abstract.

Fox R, Bourn NAD, Dennis EB, Heafield RT, Maclean IMD, Wilson RJ (2019). Opinions of citizen scientists on open access to UK butterfly and moth occurrence data. Biodiversity and Conservation, 28(12), 3321-3341. Abstract.

Dennis EB, Brereton TM, Morgan BJT, Fox R, Shortall CR, Prescott T, Foster S (2019). Trends and indicators for quantifying moth abundance and occupancy in Scotland. Journal of Insect Conservation, 23(2), 369-380. Abstract.

Banza P, Macgregor CJ, Belo ADF, Fox R, Pocock MJO, Evans DM (2019). Wildfire alters the structure and seasonal dynamics of nocturnal pollen-transport networks. FUNCTIONAL ECOLOGY, 33(10), 1882-1892. Author URL.

Bowles N, Fox R (2018). Butterflies. British Wildlife, 30(1), 53-55.

Mason SC, Palmer G, Fox R, Gillings S, Hill JK, Thomas CD, Oliver TH (2018). Corrigendum: Geographical range margins of many taxonomic groups continue to shift polewards [Biological Journal of the Linnean Society, 115, 3, (2015) (586-597)] DOI: 10.1111/bij.12574. Biological Journal of the Linnean Society, 125(2), 455-458. Abstract.

Suggitt AJ, Wilson RJ, Isaac NJB, Beale CM, Auffret AG, August T, Bennie JJ, Crick HQP, Duffield S, Fox R, et al (2018). Extinction risk from climate change is reduced by microclimatic buffering. Nature Climate Change, 8(8), 713-717.

Mason SC, Hill JK, Thomas CD, Powney GD, Fox R, Brereton T, Oliver TH (2018). Population variability in species can be deduced from opportunistic citizen science records: a case study using British butterflies. INSECT CONSERVATION AND DIVERSITY, 11(2), 131-142. Author URL.

Pearce-Higgins JW, Beale CM, Oliver TH, August TA, Carroll M, Massimino D, Ockendon N, Savage J, Wheatley CJ, Ausden MA, et al (2017). A national-scale assessment of climate change impacts on species: Assessing the balance of risks and opportunities for multiple taxa. BIOLOGICAL CONSERVATION, 213, 124-134. Author URL.

Pocock MJO, Roy HE, Fox R, Ellis WN, Botham M (2017). Citizen science and invasive alien species: Predicting the detection of the oak processionary moth <i>Thaumetopoea</i> <i>processionea</i> by moth recorders. BIOLOGICAL CONSERVATION, 208, 146-154. Author URL.

Palmer G, Platts PJ, Brereton T, Chapman JW, Dytham C, Fox R, Pearce-Higgins JW, Roy DB, Hill JK, Thomas CD, et al (2017). Climate change, climatic variation and extreme biological responses. Philos Trans R Soc Lond B Biol Sci, 372(1723). Abstract. Author URL.

Dyer RJ, Gillings S, Pywell RF, Fox R, Roy DB, Oliver TH (2017). Developing a biodiversity-based indicator for large-scale environmental assessment: a case study of proposed shale gas extraction sites in Britain. JOURNAL OF APPLIED ECOLOGY, 54(3), 872-882. Author URL.

Dennis EB, Morgan BJT, Freeman SN, Ridout MS, Brereton TM, Fox R, Powney GD, Roy DB (2017). Efficient occupancy model-fitting for extensive citizen-science data. PLOS ONE, 12(3). Author URL.

Macgregor CJ, Evans DM, Fox R, Pocock MJO (2017). The dark side of street lighting: impacts on moths and evidence for the disruption of nocturnal pollen transport. GLOBAL CHANGE BIOLOGY, 23(2), 697-707. Author URL.

Dennis EB, Morgan BJT, Brereton TM, Roy DB, Fox R (2017). Using citizen science butterfly counts to predict species population trends. CONSERVATION BIOLOGY, 31(6), 1350-1361. Author URL.

Redhead JW, Fox R, Brereton T, Olivera TH (2016). Assessing species' habitat associations from occurrence records, standardised monitoring data and expert opinion: a test with British butterflies. ECOLOGICAL INDICATORS, 62, 271-278. Author URL.

Palmer G, Hill JK, Brereton TM, Brooks DR, Chapman JW, Fox R, Oliver TH, Thomas CD (2016). Retraction of the Research Article: "Individualistic sensitivities and exposure to climate change explain variation in species' distribution and abundance changes". Sci Adv, 2(4). Abstract. Author URL.

Ovaskainen O, Roy DB, Fox R, Anderson BJ (2016). Uncovering hidden spatial structure in species communities with spatially explicit joint species distribution models. Methods in Ecology and Evolution, 7(4), 428-436. Abstract.

Dapporto L, Ciolli G, Dennis RLH, Fox R, Shreeve TG (2015). A new procedure for extrapolating turnover regionalization at mid-small spatial scales, tested on British butterflies. METHODS IN ECOLOGY AND EVOLUTION, 6(11), 1287-1297. Author URL.

Gilburn AS, Bunnefeld N, McVean Wilson J, Botham MS, Brereton TM, Fox R, Goulson D (2015). Are neonicotinoid insecticides driving declines of widespread butterflies?. PeerJ, 2015(11). Abstract.

Bowles N, Fox R (2015). Butterflies. British Wildlife, 26(6), 427-429.

Bowles N, Fox R (2015). Butterflies. British Wildlife, 26(3), 202-204.

Mason SC, Palmer G, Fox R, Gillings S, Hill JK, Thomas CD, Oliver TH (2015). Geographical range margins of many taxonomic groups continue to shift polewards. Biological Journal of the Linnean Society, 115(3), 586-597. Abstract.

Gillingham PK, Alison J, Roy DB, Fox R, Thomas CD (2015). High Abundances of Species in Protected Areas in Parts of their Geographic Distributions Colonized during a Recent Period of Climatic Change. Conservation Letters, 8(2), 97-106. Abstract.

Palmer G, Hill JK, Brereton TM, Brooks DR, Chapman JW, Fox R, Oliver TH, Thomas CD (2015). Individualistic sensitivities and exposure to climate change explain variation in species' distribution and abundance changes. Sci Adv, 1(9). Abstract. Author URL.

Suggitt AJ, Wilson RJ, August TA, Fox R, Isaac NJB, Macgregor NA, Morecroft MD, Maclean IMD (2015). Microclimate affects landscape level persistence in the British Lepidoptera. Journal of Insect Conservation, 19(2), 237-253. Abstract.

Fox R, Oliver T (2015). Much ado about Moths. Planet Earth(Spring), 26-27.

Macgregor CJ, Pocock MJO, Fox R, Evans DM (2015). Pollination by nocturnal Lepidoptera, and the effects of light pollution: a review. Ecological Entomology, 40(3), 187-198. Abstract.

Gillingham PK, Bradbury RB, Roy DB, Anderson BJ, Baxter JM, Bourn NAD, Crick HQP, Findon RA, Fox R, Franco A, et al (2015). The effectiveness of protected areas in the conservation of species with changing geographical ranges. Biological Journal of the Linnean Society Abstract.

Mair L, Hill JK, Fox R, Botham M, Brereton T, Thomas CD (2014). Abundance changes and habitat availability drive species' responses to climate change. Nature Climate Change, 4(2), 127-131. Abstract.

Bowles N, Fox R (2014). Butterflies. British Wildlife, 25(5), 358-360.

Fox R, Oliver TH, Harrower C, Parsons MS, Thomas CD, Roy DB (2014). Long-term changes to the frequency of occurrence of British moths are consistent with opposing and synergistic effects of climate and land-use changes. Journal of Applied Ecology, 51(4), 949-957. Abstract.

Suggitt AJ, Wilson RJ, August TA, Fox R, Isaac NJB, Macgregor NA, Morecroft MD, Maclean IMD (2014). Microclimate affects landscape level persistence in the British Lepidoptera. Journal of Insect Conservation Abstract.

Pagel J, Anderson BJ, O'Hara RB, Cramer W, Fox R, Jeltsch F, Roy DB, Thomas CD, Schurr FM (2014). Quantifying range-wide variation in population trends from local abundance surveys and widespread opportunistic occurrence records. Methods in Ecology and Evolution, 5(8), 751-760. Abstract.

Bowles N, Fox R (2013). Butterflies. British Wildlife, 24(6), 426-427.

Stefanescu C, Páramo F, Åkesson S, Alarcón M, Ávila A, Brereton T, Carnicer J, Cassar LF, Fox R, Heliölä J, et al (2013). Multi-generational long-distance migration of insects: Studying the painted lady butterfly in the Western Palaearctic. Ecography, 36(4), 474-486. Abstract.

Carvalheiro LG, Kunin WE, Keil P, Aguirre-Gutiérrez J, Ellis WN, Fox R, Groom Q, Hennekens S, Van Landuyt W, Maes D, et al (2013). Species richness declines and biotic homogenisation have slowed down for NW-European pollinators and plants. Ecology Letters, 16(7), 870-878. Abstract.

Fox R (2013). The decline of moths in Great Britain: a review of possible causes. Insect Conservation and Diversity, 6(1), 5-19. Abstract.

Bishop TR, Botham MS, Fox R, Leather SR, Chapman DS, Oliver TH (2013). The utility of distribution data in predicting phenology. Methods in Ecology and Evolution, 4(11), 1024-1032. Abstract.

Bowles N, Fox R (2012). Butterflies. British Wildlife, 24(1), 53-55.

Rapacciuolo G, Roy DB, Gillings S, Fox R, Walker K, Purvis A (2012). Climatic associations of British species distributions show good transferability in time but low predictive accuracy for range change. PLoS ONE, 7(7). Abstract.

Thomas CD, Gillingham PK, Bradbury RB, Roy DB, Anderson BJ, Baxter JM, Bourne NAD, Crick HQP, Findon RA, Fox R, et al (2012). Protected areas facilitate species' range expansions. Proceedings of the National Academy of Sciences of the United States of America, 109(35), 14063-14068. Abstract.

Pateman RM, Hill JK, Roy DB, Fox R, Thomas CD (2012). Temperature-dependent alterations in host use drive rapid range expansion in a butterfly. Science, 336(6084), 1028-1030. Abstract.

Mair L, Thomas CD, Anderson BJ, Fox R, Botham M, Hill JK (2012). Temporal variation in responses of species to four decades of climate warming. Global Change Biology, 18(8), 2439-2447. Abstract.

Fox R, Warren MS, Brereton TM, Roy DB, Robinson A (2011). A new Red List of British butterflies. Insect Conservation and Diversity, 4(3), 159-172. Abstract.

Asher J, Fox R, Warren MS (2011). British butterfly distributions and the 2010 target. Journal of Insect Conservation, 15(1), 291-299. Abstract.

Buckley LB, Waaser SA, MacLean HJ, Fox R (2011). Does including physiology improve species distribution model predictions of responses to recent climate change?. Ecology, 92(12), 2214-2221. Abstract.

Fox R, Randle Z, Hill L, Anders S, Wiffen L, Parsons MS (2011). Moths count: Recording moths for conservation in the UK. Journal of Insect Conservation, 15(1), 55-68. Abstract.

Fox R, Roy DB, Dennis RLH, Asher J (2010). A new look at the changing distributions of British butterflies: Different analysis, same results. Entomologist's Gazette, 61(4), 225-234. Abstract.

Fox R, Dennis RLH (2010). Winter survival of Vanessa atalanta (Linnaeus, 1758) (Lepidoptera: Nymphalidae): a new resident butterfly for Britain and Ireland?. Entomologist's Gazette, 61(2), 94-103. Abstract.

Willis SG, Hill JK, Thomas CD, Roy DB, Fox R, Blakeley DS, Huntley B (2009). Assisted colonization in a changing climate: a test‐study using two U.K. butterflies. Conservation Letters, 2(1), 46-52. Abstract.

Bowles N, Fox R (2009). Butterflies. British Wildlife, 20(3), 204-206.

Willis SG, Thomas CD, Hill JK, Collingham YC, Telfer MG, Fox R, Huntley B (2009). Dynamic distribution modelling: Predicting the present from the past. Ecography, 32(1), 5-12. Abstract.

Franco AMA, Anderson BJ, Roy DB, Gillings S, Fox R, Moilanen A, Thomas CD (2009). Surrogacy and persistence in reserve selection: Landscape prioritization for multiple taxa in Britain. Journal of Applied Ecology, 46(1), 82-91. Abstract.

Randle Z, Fox R (2008). An update on the National Moth Recording Scheme. Entomologist's Gazette, 59(2), 105-107. Abstract.

Bowles N, Fox R (2008). Butterflies. British Wildlife, 20(1), 52-54.

Bowles N, Fox R (2008). Butterflies. British Wildlife, 19(6), 434-435.

Bowles N, Fox R (2008). Butterflies. British Wildlife, 19(5), 355-357.

Bowles N, Fox R (2008). Butterflies. British Wildlife, 20(2), 125-128.

Bowles N, Fox R (2008). Butterflies. British Wildlife, 19(4), 280-281.

Fox R, Hill L, Randle Z, Anders S, Mold SA, Parsons M (2008). Moths count: the National Moth Recording Scheme. British Wildlife, 20(2), 88-96.

Bowles N, Fox R (2007). Butterflies. British Wildlife, 19(2), 127-129.

Bowles N, Fox R (2007). Butterflies. British Wildlife, 18(4), 281-282.

Bowles N, Fox R (2007). Butterflies. British Wildlife, 18(5), 356-359.

Bowles N, Fox R (2007). Butterflies. British Wildlife, 18(3), 205-207.

Bowles N, Fox R (2007). Butterflies. British Wildlife, 18(6), 433-436.

Bowles N, Fox R (2007). Butterflies. British Wildlife, 19(1), 55-57.

Menéndez R, González-Megías A, Collingham Y, Fox R, Roy DB, Ohlemüller R, Thomas CD (2007). Direct and indirect effects of climate and habitat factors on butterfly diversity. Ecology, 88(3), 605-611. Abstract.

Davies H, Brereton TM, Roy DB, Fox R (2007). Government targets for protected area management: Will threatened butterflies benefit?. Biodiversity and Conservation, 16(13), 3719-3736. Abstract.

Bowles N, Fox R (2006). Butterflies. British Wildlife, 17(5), 356-359.

Bowles N, Fox R (2006). Butterflies. British Wildlife, 17(6), 435-437.

Bowles N, Fox R (2006). Butterflies. British Wildlife, 17(4), 280-282.

Bowles N, Fox R (2006). Butterflies. British Wildlife, 17(3), 203-204.

Bowles N, Fox R (2006). Butterflies. British Wildlife, 18(2), 130-132.

Franco AMA, Hill JK, Kitschke C, Collingham YC, Roy DB, Fox R, Huntley B, Thomas CD (2006). Impacts of climate warming and habitat loss on extinctions at species' low-latitude range boundaries. Global Change Biology, 12(8), 1545-1553. Abstract.

Conrad KF, Warren MS, Fox R, Parsons MS, Woiwod IP (2006). Rapid declines of common, widespread British moths provide evidence of an insect biodiversity crisis. Biological Conservation, 132(3), 279-291. Abstract.

Menendez R, Megias AG, Hill JK, Braschler B, Willis SG, Collingham Y, Fox R, Roy DB, Thomas CD (2006). Species richness changes lag behind climate change. PROCEEDINGS OF THE ROYAL SOCIETY B-BIOLOGICAL SCIENCES, 273(1593), 1465-1470. Author URL.

Hickling R, Roy DB, Hill JK, Fox R, Thomas CD (2006). The distributions of a wide range of taxonomic groups are expanding polewards. Global Change Biology, 12(3), 450-455. Abstract.

Dennis RLH, Shreeve TG, Isaac NJB, Roy DB, Hardy PB, Fox R, Asher J (2006). The effects of visual apparency on bias in butterfly recording and monitoring. Biological Conservation, 128(4), 486-492. Abstract.

Parsons MS, Fox R, Conrad KF, Woiwod IP, Warren MS (2005). British moths: Throwing light on a new conservation challenge. British Wildlife, 16(6), 386-394.

Bowles N, Fox R (2005). Butterflies. British Wildlife, 16(5), 357-359.

Bowles N, Fox R (2005). Butterflies. British Wildlife, 16(4), 277-279.

Bowles N, Fox R (2005). Butterflies. British Wildlife, 16(3), 205-207.

Bowles N, Fox R (2005). Butterflies. British Wildlife, 17(1), 51-53.

Bowles N, Fox R (2005). Butterflies. British Wildlife, 16(6), 432-434.

Bowles N, Fox R (2005). Butterflies. British Wildlife, 17(2), 127-128.

Moilanen A, Franco AMA, Early RI, Fox R, Wintle B, Thomas CD (2005). Prioritizing multiple-use landscapes for conservation: Methods for large multi-species planning problems. Proceedings of the Royal Society B: Biological Sciences, 272(1575), 1885-1891. Abstract.

Bowles N, Fox R (2004). Butterflies. British Wildlife, 15(5), 359-361.

Bowles N, Fox R (2004). Butterflies. British Wildlife, 16(1), 56-58.

Bowles N, Fox R (2004). Butterflies. British Wildlife, 16(2), 130-131.

Bowles N, Fox R (2004). Butterflies. British Wildlife, 15(3), 210-212.

Bowles N, Fox R (2004). Butterflies. British Wildlife, 15(4), 277-279.

Thomas JA, Telfer MG, Roy DB, Preston CD, Greenwood JJD, Asher J, Fox R, Clarke RT, Lawton JH (2004). Comparative Losses of British Butterflies, Birds, and Plants and the Global Extinction Crisis. Science, 303(5665), 1879-1881. Abstract.

Conrad KF, Woiwod IP, Parsons M, Fox R, Warren MS (2004). Long-term population trends in widespread British moths. Journal of Insect Conservation, 8(2-3), 119-136. Abstract.

Conrad KF, Woiwod IP, Parsons M, Fox R, Warren MS (2004). Long-term population trends in widespread British moths. Journal of Insect Conservation, 8(2-3), 119-136.

RJWilson, Fox R, Roy DB, Thomas CD (2004). Spatial patterns in species distributions reveal biodiversity change. Nature, 432(7015), 393-396.

Bowles N, Fox R (2003). Butterflies. British Wildlife, 14(6), 437-440.

Bowles N, Fox R (2003). Butterflies. British Wildlife, 15(1), 58-60.

Bowles N, Fox R (2003). Butterflies. British Wildlife, 14(5), 359-360.

Bowles N, Fox R (2003). Butterflies. British Wildlife, 15(2), 126-128.

Bowles N, Fox R (2003). Butterflies. British Wildlife, 14(3), 209-210.

Bowles N, Fox R (2003). Butterflies. British Wildlife, 14(4), 284-285.

Hill JK, Fox R (2003). Climate change and British butterfly distributions. Biologist, 50(3), 106-110. Abstract.

Bowles N, Fox R (2002). Butterflies. British Wildlife, 13(6), 436-438.

Bowles N, Fox R (2002). Butterflies. British Wildlife, 14(1), 56-58.

Bowles N, Fox R (2002). Butterflies. British Wildlife, 13(5), 360-361.

Bowles N, Fox R (2002). Butterflies. British Wildlife, 14(2), 132-133.

Hill JK, Thomas CD, Fox R, Telfer MG, Willis SG, Asher J, Huntley B (2002). Responses of butterflies to twentieth century climate warming: Implications for future ranges. Proceedings of the Royal Society B: Biological Sciences, 269(1505), 2163-2171. Abstract.

Hill JK, Collingham YC, Thomas CD, Blakeley DS, Fox R, Moss D, Huntley B (2001). Impacts of landscape structure on butterfly range expansion. Ecology Letters, 4(4), 313-321. Abstract.

Warren MS, Hill JK, Thomas JA, Asher J, Fox R, Huntley B, Roy DB, Telfer MG, Jeffcoate S, Harding P, et al (2001). Rapid responses of British butterflies to opposing forces of climate and habitat change. Nature, 414(6859), 65-69. Abstract.

Fox R, Warren M, Harding P, Asher J, Jeffcoate G, Jeffcoate S (2001). The millennium atlas of butterflies in Britain and Ireland. British Wildlife, 12(3), 173-178.

Fox R (1998). Butterflies for the new millennium - a new atlas of butterflies in Britain and Ireland. British Wildlife, 9(3), 176-179.

Chapters

Fox R, Conrad KF, Parsons MS, Warren MS, Woiwod IP (2010). Moths. In (Ed) Silent Summer: the State of Wildlife in Britain and Ireland, 448-470. Abstract.

Conrad KF, Fox R, Woiwod IP (2007). Monitoring biodiversity: Measuring long-term changes in insect abundance. In (Ed) Insect Conservation Biology: Proceedings of the Royal Entomological Society's 23rd Symposium, 203-225.

Warren MS, Bourn N, Brereton T, Fox R, Middlebrook I, Parsons MS (2007). What have red lists done for us? the values and limitations of protected species listing for invertebrates. In (Ed) Insect Conservation Biology: Proceedings of the Royal Entomological Society's 23rd Symposium, 76-91.

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